The clonal origin of multiple lung tumors is uncertain.

多发肺肿瘤的起源克隆尚不清楚。

They arise when there is a clonal expansion of a mutated cell.

一个突变细胞如发生克隆性扩增，就引起这些大量突变。

Clonal anergy and suppressor cells are main mechanisms of the tolerance.

克隆不应答和抑制细胞是耐受的主要机制。

Clonal propagation can also give some large gains in a relatively short time.

繁殖无性系也可以在较短期间内提供较大的增益。

And then the clonal selection algorithm using this clonal operator is discussed.

并对应用此算子的克隆选择算法进行了讨论。